In 2003, the Human Genome Project's ambitious goals had all been met or surpassed. The sequences produced by the Human Genome Project covered about 99 percent of the human genome's gene-containing regions. Not only was the project finished two-and-a-half years ahead of time, but it was also significantly under budget. In addition, to help researchers better understand the meaning of the human genetic instruction book, the project successfully undertook a wide range of other goals: from sequencing the genomes of organisms used in disease research, to developing new technologies for studying whole genomes. The Human Genome Project has been compared to the moon-landing project as an outstanding scientific achievement of humankind.